KEOGH RIVER
Long-term Research Watershed with Anadromous Salmonids

KEOGH RIVER SCIENCE PUBLICATIONS
Bailey, C.J., and J.W. Moore. 2020. Resource pulses increase the diversity of successful competitors in a multi-species stream fish assemblage. Ecosphere. 11(9): e03211. doi: 10.1002/ecs2.3211. (PDF).
Bailey, C.J., L.C. Andersson, M. Arbeider, K. Bradford, and J.W. Moore. 2019. Salmon egg subsidies and interference competition among stream fishes. Environmental Biology of Fishes. 1-12. doi: 10.1007/s10641-019-00880-9 (PDF).
Bailey, C. J., D. C. Braun, D. McCubbing, J. D. Reynolds, B. Ward, T. D. Davies, and J. W. Moore. 2018. The roles of extrinsic and intrinsic factors in the freshwater life-history dynamics of a migratory salmonid. Ecosphere 00(00): e02397. 10.1002/ecs2.2397 (link)
Atlas, W. I., T. W. Buehrens, D. J. F. Mccubbing, R. Bison, and J. W. Moore. 2015. Implications of spatial contraction for density dependence and conservation in a depressed population of anadromous fish. Canadian Journal of Fisheries & Aquatic Sciences 12:1–12. PDF
Friedland, K. D., B. R. Ward, D. W. Welch, and S. A. Hayes. 2014. Postsmolt growth and thermal regime define the marine survival of steelhead from the Keogh River, British Columbia. Marine and Coastal Fisheries: Dynamics, Management, and Ecosystem Science 6:1–11. PDF
Melnychuk, M. C., Welch, D. W. & Walters, C. J. 2010. Spatio-temporal migration patterns of Pacific salmon smolts in rivers and coastal marine waters. PLoS ONE 5. PDF
Melnychuk, M. C. 2009. Estimation of survival and detection probabilities for multiple tagged salmon stocks with nested migration routes, using a large-scale telemetry array. Marine and Freshwater Research 60, 1231–1243. PDF
Halpenny, C. & Gross, M. R. 2008. Macro- and microparasite infection profiles of hatchery fish before and after release from a conservation hatchery (steelhead trout, Oncorhynchus mykiss). Écoscience 15, 417–422. PDF
Ward, B. R., McCubbing, D. J. F. & Slaney, P. A. 2007. Stream restoration for anadromous Salmonids by the addition of habitat and nutrients. In Salmon at the edge (Mills, D., ed), pp. 33–254 Wiley-Blackwell.
Ward, B. R., P. A. Slaney, and D. J. F. McCubbing. 2006. Watershed restoration to reconcile fisheries and habitat impacts at the Keogh River in coastal British Columbia. American Fisheries Society Symposium:587–602. PDF
Walters, J. C. 2005. Conservation value of alternative life-history tactics in steelhead trout (Oncorhynchus mykiss). University of Toronto. PDF
Welch, D. W., Boehlert, G. W. & Ward, B. R. 2003. POST–the Pacific Ocean salmon tracking project. Oceanologica Acta 25, 243–253. PDF
Smith, B. D., Ward, B. R. & Welch, D. W. 2000. Trends in wild adult steelhead (Oncorhynchus mykiss) abundance in British Columbia as indexed by angler success. Canadian Journal of Fisheries and Aquatic Sciences 57, 225–270. PDF
Smith, B. D., and B. R. Ward. 2000. Trends in wild adult steelhead (Oncorhynchus mykiss) abundance for snowmelt-driven watersheds of British Columbia in relation to freshwater discharge. Canadian Journal of Fisheries and Aquatic Sciences 57:271–284. PDF
Welch, D. W., Ward, B. R., Smith, B. D. & Eveson, J. P. 2000. Temporal and spatial responses of British Columbia steelhead (Oncorhynchus mykiss) populations to ocean climate shifts. Fisheries Oceanography 9, 17–32. PDF
Ward, B. R. 2000. Declivity in steelhead (Oncorhynchus mykiss) recruitment at the Keogh River over the past decade. Canadian Journal of Fisheries and Aquatic Sciences 57:298–306. PDF
Walters, C. & Ward, B. 1998. Is solar radiation responsible for declines in marine survival rates of anadromous salmonids that rear in small streams? Canadian Journal of Fisheries and Aquatic Sciences 55, 2533–2538. PDF
Welch, D. W., Ward, B. R., Smith, B. D. & Eveson, J. P. 1998. Influence of the 1990 ocean climate shift on British Columbia steelhead (Oncorhynchus mykiss) and coho (O. kisutch) populations. In Proceedings ‘Aha Huliko’a Hawaiian Winter Workshop, Biotic Impacts of Extratropical Climate Variability in the Pacific (Holloway, G., Muller, P., Henderson, D., eds), pp. 77–87 University of Hawaii at Manoa. PDF
Taylor, E. B. 1995. Genetic variation at minisatellite DNA loci among North Pacific populations of steelhead and rainbow trout (Oncorhynchus mykiss). Journal of Heredity1 86, 354–363. PDF
Ward, B. R., and P. A. Slaney. 1993. Egg-to-smolt survival and fry-to-smolt density dependence of Keogh River steelhead trout. Canadian Journal of Fisheries & Aquatic Sciences 118:209–217. PDF
Irvine, J. R. & Johnston, N. T. 1992. Coho salmon (Oncorhynchus kisutch) use of lakes and streams in the Keogh River Drainage, British Columbia. Northwest Science 66, 15–25. PDF
Irvine, J. R., Ward, B. R., Teti, P. A. & Cousens, N. B. F. 1991. Evaluation of a method to count and measure live salmonids in the field with a video camera and computer. North American Journal of Fisheries Management 11, 20–26. PDF
Johnston, N. T., C. J. Perrin, P. A. Slaney, and B. R. Ward. 1990. Increased juvenile salmonid growth by whole-river fertilization. Canadian Journal of Fisheries and Aquatic Sciences 47:862–872. PDF
Ward, B. R. & Slaney, P. A. 1990. Returns of pen-reared steelhead from riverine, estuarine, and marine releases. Transactions of the American Fisheries Society 119, 492–499. PDF
Irvine, J. R. & Ward, B. R. 1989. Patterns of timing and size of wild coho salmon (Oncorhynchus kisutch) smolts migrating from the Keogh River Watershed on Northern Vancouver Island. Canadian Journal of Fisheries & Aquatic Sciences 46, 1086–1094. PDF
Ward, B. R., P. A. Slaney, A. R. Facchin, and R. W. Land. 1989. Size-biased survival in steelhead trout (Oncorhynchus mykiss): back-calculated lengths from adults’ scales compared to migrating smolts at the Keogh River, British Columbia. Canadian Journal of Fisheries & Aquatic Sciences 46:1853–1858. PDF
Swales, S., Caron, F., Irvine, J. R. & Levings, C. D. 1988. Overwintering habitats of coho salmon (Oncorhynchus kisutch) and other juvenile salmonids in the Keogh River system, British Columbia. Can.J.Zool. 66, 254–261. PDF
Ward, B. R., and P. A. Slaney. 1988. Life history and smolt-to-adult survival of Keogh River steelhead trout (Salmo gairdneri) and the relationship to smolt size. Canadian Journal of Fisheries and Aquatic Sciences 45:1110–1122. PDF
Beacham, T. D. 1985. Meristic and morphometric variation in pink salmon (Oncorhynchus gorbuscha) in southern British Columbia and Puget Sound. Canadian Journal of Zoology 63, 366–372. PDF
Perrin, C. J., M. L. Bothwell, and P. A. Slaney. 1987. Experimental enrichment of a coastal stream in British Columbia: effects of organic and inorganic additions on autotrophic periphyton production. Canadian Journal of Fisheries and Aquatic Sciences 44:1247–1256. PDF
Qadri, S. U. 1959. Some morphological differences between the subspecies of cutthroat trout, Salmo clarki clarki and Salmo clarki lewisi, in British Columbia. Journal of the Fisheries Research Board of Canada 16, 903–922. PDF
KEOGH RIVER GOVERNMENT REPORTS
J.M.S. Harding. 2016. Adult steelhead trout (Oncorhynchus mykiss) and salmonid smolt migrations at the Keogh River, BC, during winter 2015 and spring 2016. BC Ministry of Forests, Lands and Natural Resource Operations, Victoria, BC. PDF
Ward, B. R. 2011. A science review on policy for augmentation and supplementation of wild steelhead trout with hatchery fish in British Columbia. Victoria, BC. PDF
Ward, B. R. 2006. The case for wild steelhead recovery without artificial fish culture intervention. Vancouver, BC. PDF
Korman, J., Higgins, P. S. & Melville, C. C. 2005. Incorporating the effects of migratory behaviour and river conditions into estimates of steelhead escapement. 56. PDF
McCubbing, D. J. F. 2002. Adult steelhead trout and salmonid smolt migration at the Keogh River, B.C. during spring 2002. Port Hardy, BC. PDF
McCubbing, D. J. F., and B. R. Ward. 2001. Stream rehabilitation in British Columbia’s watershed restoration program: juvenile salmonids in the Keogh River compared to the untreated Waukwaas River in 2001. Vancouver, BC. PDF
McCubbing, D. J. F., B. R. Ward, and L. Burroughs. 2000. Salmonid escapement enumeration on the Keogh River: a demonstration of a resistivity counter in British Columbia. Vancouver, BC. PDF
McCubbing, D. J. F., and B. R. Ward. 1999. Stream rehabilitation in British Columbia’s watershed restoration program: positive response by juvenile salmonids in the Keogh River compared to the untreated Waukwaas River in 1999. Vancouver, BC. PDF
Slaney, P. A., and K. I. Ashley. 1999. Case studies of whole-stream fertilization in British Columbia. Victoria, BC. PDF
McCubbing, D. J. F., and B. R. Ward. 1998. Stream rehabilitation in British Columbia’s watershed restoration program: juvenile salmonid response in the Keogh and Waukwaas Rivers 1998. Page Fisheries Research. Vancouver, BC. PDF
Ward, B. R., and D. J. F. McCubbing. 1998. Adult steelhead trout and salmonid smolts at the Keogh River during spring 1998 and comparison to the historic record. Page Fisheries Technical Circular. Vancouver, BC. PDF
Slaney, P. A., and D. Zaldokas. 1997. Fish habitat rehabilitation procedures. Page (P. A. Slaney and D. Zaldokas, Eds.) Watershed Restoration Technical Circular. Vancouver, BC. PDF
McCubbing, D. J. F., and B. R. Ward. 1997. The Keogh and Waukwaas Rivers paired watershed study for B.C.’s watershed restoration program: juvenile salmonid enumeration and growth 1997. Page Watershed Restoration Project Report No. 6. Vancouver, BC. PDF
Ashley, K. & Mouldey, S. 1996. Restoring stream productivity with Nutri-Stones. B.C.’s Stream Restoration Technical Bulletin 5–9. PDF
Slaney, P. A., Ward, B. R. & Berg, L. 1990. A preliminary assessment of the effect of external abrasion on the smolt-to-adult survival of net-pen cultured steelhead trout. Fisheries Technical Circular 88, 1–11. PDF
Ward, B. R., J. A. Burrows, and D. L. Quamme. 1990. Adult steelhead population size and salmonid migrants of the Keogh River during the spring, 1990. Vancouver, BC. PDF
Ward, B. R., and J. C. Wightman. 1989. Monitoring steelhead trout at the Keogh River as an index of stock status and smolt-to-adult survival: correlations with other data sources. Vancouver, BC. PDF
Ward, B. R. 1988. Trends and forecasts from winter -run steelhead at the Keogh River. Vancouver, BC. PDF
Lirette, M. G., Hooton, R. S. & Lewynksky, V. A. 1987. Preliminary steelhead production capability estimates for selected Vancouver Island streams. Fisheries Techni 74, 24 p. PDF
Johnston, N. T., J. R. Irvine, and C. J. Perrin. 1986. A comparitive evaluation of fence count, mark-recapture and Bendix sonar estimates of salmon escapements in the Keogh River, a variable-flow coastal B.C. stream. West Vancouver, BC. PDF
Slaney, P. A., Billings, S. J. & Smith, H. A. 1980. Experimental stocking of steelhead fry above barriers in a high gradient tributary of the Keogh River: progress 1978 and 1979. Fisheries Technical Circular 48, 28 p. PDF
Smith, H. A., and P. A. Slaney. 1980. Age, growth, survival and habitat of anadromous Dolly Varden (Salvelinus malma) in the Keogh River, British Columbia. Page Fisheries Management Report No. 76. Victoria, BC. PDF
Ward, B. R. & Slaney, P. A. 1979. Evaluation of in-stream enhancement structures for the production of juvenile steelhead trout and Coho salmon in the Keogh River: progress 1977-1978. Vol. 45. PDF
Mottram, W. 1977. Design and construction of the Keogh River fish fence. Page Fisheries Technical Circular. Victoria, BC. PDF
Fraser, F. J., D. T. Lightly, and D. D. Bailey. 1974. An inventory of east coast Vancouver Island streams important to Chum Salmon. PDF